Antwi, E. K. et al. Paper: Land Use and Landscape Structural Changes in the Ecoregions of Ghana Effah Kwabena Antwi∗1, John Boakye-Danquah∗2, Stephen Boahen Asabere∗3, Gerald A. B. Yiran∗4, Seyram Kofi Loh∗4, Kwabena Gyekye Awere∗4, Felix K. Abagale∗5, Kwabena Owusu Asubonteng∗6, Emmanuel Morgan Attua∗7, and Alex Barimah Owusu∗7 ∗1Integrated Research System for Sustainability Science (IR3S), the University of Tokyo 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-8654, Japan E-mail: antwi@unu.edu ∗2Department of Geography and Resource, University of Ghana, Ghana ∗3Department of Geosceinces, Technische Universitaet Dresden, Germany ∗4Department of Geography and Resource Development, University of Ghana, Ghana ∗5Faculty of Agriculture, University for Development Studies, Tamale, Ghana ∗6United Nations University – Institute for Natural Resources in Africa ∗7Department of Geography and Resource Development, University of Ghana [Received February 18, 2014; accepted July 23, 2014] In recent years, land use (LU) and landscape structure in ecoregions around the world have been faced with enormous pressures, from rapid population growth to urban sprawl. A preliminary account of changes in land cover (LC) and landscape structure in the ecore- gions of Ghana is missing from the academic and re- search literature. The study therefore provides a pre- liminary assessment of the changing LU and landscape structure in the ecoregions of Ghana, identifying the causes and assessing their impact on land-based re- sources, and on urban and agricultural development. LU/LC maps produced from 30 m resolution Landsat TM5 in 1990 and ETM+ in 2000 were classified into dominant land cover types (LCTs) and used to sur- vey the changing landscape of Ghana. LC-change map preparation was done with change detection extension “Vera¨nderung” (v3) in an ArcGIS 10.1 environment. At the class level, Patch Analyst version 5.1 was used to calculate land use (LU) statistics and to provide land- scape metrics for LU maps extracted from the satel- lite imagery. The results showed that commonly ob- served LCCs in the ecoregions of Ghana include con- version of natural forest land to various forms of cul- tivated lands, settlements, and open land, particularly in closed and open forest and savannah woodland. The dominant LU types in the ecoregions of Ghana are arable lands, which increased by 6168.98 km2 . For- est and plantation LCTs decreased in area and were replaced by agricultural land, forest garden, and open land. Afforestation rarely occurred except in the rain- forests. The mean patch size (MPS), a measure of frag- mentation, was generally reduced consistently from 1990 to 2000 in all the ecoregions. Similar results that indicated increased fragmentation were an increased number of patches (NumP) and the Shannon diver- sity index (SDI). Habitat shape complexity inferred from mean shape index (MSI) decreased in all ecore- gions except for rainforest and wet evergreen. The SDI and Shannon evenness index (SEI) showed that habi- tat diversity was highest in the coastal savannah and the deciduous forest ecoregions. The main drivers of changes in the LUs and landscape structure are de- mand for land and land-based natural resources to support competing livelihoods and developmental ac- tivities in the different ecoregions. Keywords: landscape metrics, land change, landscape planning, urban sprawl, fragmentation 1. Introduction Ecological regions, also known as ecoregions across the globe, vary in composition and structure. As a land resource base, each ecoregion is gradually transformed through human use and management, and environmen- tal processes. In the twentieth century, variations in cli- mate and increased land use (LU) intensity have acceler- ated changes in landscape composition and configuration in the world’s ecoregions. For example, the conversion of 13 million ha of forest land to other land use types (LUTs) globally and the loss of associated ecosystem services are major concerns [1]. More than 26% of such changes have occurred in Africa. The high rate of forest cover loss in Africa can be attributed to overdependence on land-based resources, as about 70% of the population in Africa uses the forest as the principal source of income or food (World Bank, 2006). In Ghana, the deforestation rate stands at 65,000 km2 annually, and it has been projected that all forests will be depleted in 25 years if the current rate of deforestation continues [2]. Ghana is divided into seven major ecoregions (Fig. 1), based mainly on the differences in climate (rainfall) and soils [3, 4]. Ecoregions and their characteristic climate and soil types affect human well-being and environmental 452 Journal of Disaster ResearchVol.9 No.4, 2014 Land Use and Landscape Structural Changes in the Ecoregions of Ghana processes through the distribution of natural resources for diverse socioeconomic needs. These regions have highly significant spatial and temporal implications for the sus- tainable management of landscape structure and compo- sition. In terms of the land cover types (LCTs) in the ecore- gions of Ghana, human and environmental activities have led to the development of patches on the land surface. These patches are in locations that serve as ecosystems having socio-economic, social, and ecological signifi- cance for the welfare of the surrounding communities. Patch dynamics depict the changes in the spatial pat- terns of the landscapes. The ecological and environmen- tal processes that produce these patterns, as well as the internal dynamics of changes in these patches over time, are essential concepts in landscape ecology. The spatial and temporal properties of patch dynamics help offer the dimensions of space and time to managing both natural and semi-natural landscapes, their resource composition, and conservation of other complex systems [6]. Patches are created when portions of the vegetation cover are removed and/or changed to other covers, even- tually resulting in a permanent land cover change (LCC). “Land cover is an observed physical cover, including nat- ural or planted vegetation and human structures (trans- portation networks, buildings etc.) that covers the earth’s surface” [5]. LCC can occur naturally; however, it is mostly driven by human-environment interactions, such as LU systems and climate change [6]. Therefore LC changes differ significantly from region to region and from community to community because of differences in interactions among the natural, social, economic, and po- litical factors in a particular area [7]. In Ghana differ- ences in LU, deforestation/devegetation, and urbanization are some of the human-induced observable changes tak- ing place countrywide, along with an increase in the con- struction of dams/dugouts in the interior savannah [8]. All of these contribute to landscape fragmentation. The Ghanaian landscape structure is heterogeneous with a characteristic combination of natural environment and human activities [9]. As in other agro-ecological zones, the breaking up of the landscape into smaller patches owing to human and natural forces [10] is the main transformative agent. Anthropogenic activities, such as agriculture and urban development, are considered the origin of habitat fragmentation and heterogeneity [11]. For example, the rainforest in Ghana is believed to have decreased by 17.9% from 1975 to 2000; yet, most studies on landscape structure and land use change (LUC) have focused largely on sub-regional levels, making it difficult to apply the outcomes of such assessments to national- level decision making [11]. Recent developments in technology, especially such spatial technologies as GIS and remote sensing, offer essential tools for studying the ecology of large land- scapes [12, 13] more quickly and at frequent intervals. These tools have facilitated the capturing of the spatial and temporal characteristics of patches in a landscape. Though spatial and temporal assessment of LUCs and Fig. 1. regional map of Ghana showing the seven ecoregions of Ghana. quantification of landscape structures are not new in land- scape ecology, a primary account of these changes in the ecoregions of Ghana is missing in the academic and re- search literature. This paper provides an analysis of changes in LU and landscape structure from 1990 to 2000 in the ecoregions of Ghana. Specifically, it seeks to identify causes for the changes and their impact on water resources and on for- est, urban, and agricultural development. The research is guided by the following questions: (a) What are the ma- jor LUTs in the ecoregions of Ghana? (b) What are the impacts of LU and landscape structural changes such as forest loss, agricultural expansion, and urban sprawl? 2. Methodologies 2.1. Study Area Ghana is located between latitudes 4◦44′N and 11◦11′N and longitudes 3◦11′W and 1◦11′E (Fig. 1) with a total landmass of about 239,150 km2. Fig. 1 shows the seven ecoregions of Ghana on a regional map. The population of Ghana has been increasing steadily at an average in- tercensal growth rate of 2.5% and now stands at about 25 million [23]. Agriculture is a dominant part of Ghana’s economy, accounting for 35% of the country’s GDP. Journal of Disaster ResearchVol.9 No.4, 2014 453 Antwi, E. K. et al. Ghana’s climate varies from the tropical unimodal monsoon type in the north to the bimodal equatorial type in the south [3]. In general, rainfall increases from south to north and is a main moisture source for the agricul- tural enterprises in the country [3]. Mean annual rain- fall varies across the seven ecoregions, with wet ever- green forest recording the highest mean annual rainfall of above 2200 mm followed by rainforest (2200 mm), de- ciduous forest (1500 mm), transitional zone (1300 mm), Guinea savannah (1100 mm) Sudan savannah (1000 mm) and coastal savannah (800 mm) [3]. The main natural hazards in Ghana are drought, flood, and landslides. The dry dusty northeastern har- mattan winds blow across the country from January to March [13, 14]. The main soil texture classifications in Ghana are sandy loam and loam, though coarse sandy and clay loams can be found at the lower latitudes. The main soil types are Alfisols, Plinth Luvisols, and their integrates [15]. Soils in the south (forest zones) of Ghana are grouped under Oxisols, Ochrosols, Acid Gleysols and Lateritic. The soils in the south are porous, well drained, and gener- ally loamy compared to the soils of the north (savannah zones), which are poor in nutrients and heavily depen- dent on humus and fertilizer [16]. Approximately 47% of the soils in the northern savannah zones are unsuitable for crop production, with 25% being marginal and only 28% suitable. The three savannah regions in the northern part are covered by savannah grassland with bands of drought- resistant trees, such as baobab (Adansonia digitata), dawadawa (Parkia biglobosa), Shea (Vitellaria para- doxa), neem (Azadiracta indica) and acacia (Acacia Nilotica) at varying densities. The southern part of the country has dominantly evergreen and semi-deciduous forests, consisting mainly of such tropical hardwood trees as mahogany, odum, ebony, silk cotton and kolas. 2.2. Data Sources and Spatial Data Processing The satellite images used for the study came from Landsat TM5 data acquired in 1990 and ETM+ in 2000. The entire country is covered by 16 scenes with path/row ranging from 192-056 in the southeast to 195-052 in the Northwest. It must be mentioned that the available Land- sat scenes after the year 2000 have many failed scan lines (black lines) with no data, which rendered them unsuit- able for classification; therefore, they could not be used for the analyses. The spatial resolution of the Landsat data was 30m*30m spatial resolution. The images used were cho- sen in the vegetation period–thus, between the months of May and September. The search criteria were set for im- ages with less than 10% cloud cover. Some images un- avoidably had some clouds, though these areas were cor- rected through the use of historical data to replace the cloud-covered parts. Satellite data was processed with ERDAS Imagine 2013. The 16 scene images were first mosaicked to form a single image of the entire country before the classification was done. Table 1. Producer accuracy and user accuracy of classifica- tion output for all land user types in the ecoregions. Land cover types 1990 2000 Producer accuracy User accuracy Producer accuracy User accuracy Agricultural land 93 92 88 92 Built-up areas 87 81 94 92 Closed forest 89 95 80 91 Forest garden 92 90 97 95 Grassland with trees 88 86 87 93 Open forest 91 94 89 91 Open land 91 91 80 86 Plantation 91 96 85 96 Savannah woodland 90 91 94 93 Shrubland and thicket 92 92 88 93 Water body 90 96 96 90 Wetland 92 95 94 91 2.3. Image Classification, Change Detection, and Classification Accuracy Issues The mosaicked images of the ecoregions for the re- spective years were classified into 12 LCTs (Table 1), using supervised classification with the maximum like- lihood classification algorithm in ERDAS Imagine soft- ware. During the classification, clouds were classified as a separate LCT and later merged with the appropriate class of the main LCTs. Landsat TM5 and ETM+ imagery for the entire Ghana landscape was collected and merged for the years 1990 and 2000 to analyze the changing land- scape. Overall, 2,750 sample points were collected for 1990 and 2,610 sample points were used for 2000 by employ- ment of a simple random sampling technique. These sam- pled points were divided into two parts: the first was used to generate spectral signatures for the classification and the second as ground truth for an accuracy assessment. An overall accuracy of 92% and 92.5%was ensured in the final classification output for the 1990 and 2000 images, respectively [17]. Overall Kappa statistics in the classi- fication output for 1990 and 2000 images are 91.00 and 82.7% respectively. The producer and user accuracies of classifications in all land user types in the ecoregions of Ghana are shown in Table 1. In each ecoregion, the dominant vegetation types or LC features were used to represent the LU classes [18]. In all, 12 classes were identified for the whole country. Table 3 shows the major land use types or classes in the ecoregions of Ghana. The classification of land use types in each ecoregion was followed by a detection of changes in the LUs between the two years. The extension “Vera¨nderung” (v3) aided quantification of changes from LCT in 1990 to other LCTs in 2000 [18]. The LUC maps that were produced showed three classes: reduction in LC classes (negative change), no change, and increases in LC classes (positive change) between the two periods. 454 Journal of Disaster ResearchVol.9 No.4, 2014 Land Use and Landscape Structural Changes in the Ecoregions of Ghana Table 2. Landscape metrics descriptions and their indicator ranges use in the study of ecological features. Ecological feature Metrics indicators Description Range Habitat richness /No. of patches fragmentation Number of Patches (NumP) It is a measure of the extent of subdivision or fragmentation of the habitat type. NumP = 1 when the landscape or class consists of a single patch NumP ≥ 1 without limit Edge density (ED) It measures habitat length in a landscape. ED = 0 when the entire landscape and landscape border, if present, consists of the corre- sponding patch type. ED ≥ 0 without limit Patch/habitat size Mean patch size (MPS) The range in MPS is limited by the grain and extent of the image and the minimum patch size in the same manner as patch area. MPS > 0 Mean shape index (MSI) It measures the average patch shape or perimeter-to-area ratio, for a patch type or patches in the landscape. MSI = 1 when all patches of the corresponding patch type are cir- cular (vector). It increases without limit as the patch shapes become more irregular MSI ≥ 1 without limit Evenness habitat heterogeneity Shannon evenness index (SEI) It measures distribution of area among patch types SEI = 0 when the landscape contains only 1 patch (i.e., no diver- sity) and approaches 0 as the distribution of area among the different patch types becomes increasingly uneven. SEI =1 when distribu- tion of area among patch types is perfectly even (i.e., proportional abundances are the same). 0 ≤ SEI ≤ 1 Habitat diversity Shannon diversity index (SDI) It is a measure of diversity in community ecology SDI= 0 when the landscape contains only 1 patch (i.e., no diversity). SDI increases as the number of different patch types (patch richness, PR) increases. SDI ≥ 0 without limit 2.4. Landscape Structure Analyses The thematic LC maps from 1990 and 2000 were sub- jected to landscape statistics estimations using Patch An- alyst version 5.1. Patch Analyst offers a comprehensive choice of landscape metrics at the landscape level [18]. Representative metrics were selected for land habitat shape complexity (mean shape index, edge density, land- scape shape index) fragmentation (core area metric e.g. number of patches, mean patch size), landscape compo- sition and diversity (Shannon diversity index, Shannon evenness index). The metrics were selected based on their comparability, robustness and sensitivities to spatio- temporal landscapes [1, 10, 19]. Table 2 gives descrip- tions of metrics used. Most metrics at this level describe similar characteristics in the landscape structure; there- fore, further selection of landscape metrics was done [19] to exclude highly correlated metrics using correlation analysis with acceptable multicollinearlity threshold (r = 0.8). Garbarino et al. (2013) explained how to exclude highly correlated metrics and landscape metrics selection. 3. Results 3.1. Land Cover Distribution in the Various Ecore- gions of Ghana First, the study presents the major LCTs in Ghana. In both study periods, 12 LCTs were dominant in the seven ecoregions. A description of these LCTs is provided in Table 3. 3.1.1. Distribution of Agricultural Land and Forest Garden The dominant land-use types in the ecoregions of Ghana are agricultural land and forest garden. These two constitute the arable land and are found in all the ecore- gions of Ghana. Agricultural land generally decreased in all the ecoregions of Ghana between 1990 and 2000 ex- cept in the Guinea savannah, coastal savannah and de- ciduous forests in 2000 (Table 4). Comparing differ- ent ecoregions, the highest decrease in agricultural land occurred in the transition zone. Conversely, two ecore- gions recorded a positive change in agricultural lands. Of the two ecoregions that recorded increases in agricultural lands, the coastal savannah recorded the highest. There was a substantial net increase in the forest gar- den during the study period. Except for Sudan savannah, deciduous forest, and coastal savannah, all other ecore- gions had an increase in forest garden. The highest in- crease was found in the transition zone, with Guinea sa- vannah and rainforest recording largely similar hectarage increases. The decrease in forest garden in the decidu- ous forest was the highest in all ecoregions, followed by coastal savannah and Sudan savannah. The increase in forest gardenwas largely taken from sa- vannah woodlands in the Guinea savannah, Sudan savan- nah, and transition zones. Others were taken from open forest, closed forest, and plantation in the wet evergreen, deciduous forest, and rainforest zones; and from grassland with trees in the transition ecoregion. Forest garden also increased in the coastal savannah (Fig. 2). The decreased area in agricultural land was replaced mainly with savan- Journal of Disaster ResearchVol.9 No.4, 2014 455 Antwi, E. K. et al. T ab le 3. C om po ne nt of m aj or L U /L C ty pe s in th e ec or eg io ns of G ha na . L U /L C Ty pe D es cr ip tio n of L U /L C Ty pe s A gr ic ul tu ra l la nd A re as w he re ov er 50 % is un de r ag ri cu ltu re ,e xc lu di ng tr ee cr op s. In cl ud es gr az in g la nd s. Fo re st ga rd en Fo re st ga rd en in G ha na of te n ha s a m ix tu re of fa rm la nd s in na tu ra lly or se m i- na tu ra lg ro w n fo re st . T hi s ca te go ry is us ed fo r ar ea s w he re it is di ffi cu lt to se pa ra te fo re st an d ag ri cu ltu ra ll an d. Pl an ta tio n M ai nl y tr ee cr op pl an ta tio ns of co co a an d oi lp al m ;m ay in cl ud e fo re st pl an ta tio ns . C lo se d fo re st H as a hi gh de ns ity of fo re st tr ee s, m os tly pr ot ec te d, w ith a ca no py co ve r of > 60 % . It in cl ud es tr ad iti on al gr ov es ,c lo se d fo re st pl an ta tio ns ,f or es t, an d w ild lif e pr ot ec te d ar ea s. O pe n fo re st H ig h de ns ity of fo re st tr ee s; m ay be un de r pr ot ec tio n bu t m os tly is no t, w ith a ca no py co ve r of < 60 % . N or m al ly is de gr ad ed cl os ed fo re st m ad e of fo re st pl an ta tio ns an d in cl ud in g a fe w pa tc he s of fa rm la nd s. G ra ss la nd w ith /w ith ou tt re es Sa va nn ah la nd ch ar ac te ri ze d by gr as se s w ith sp ar se di st ri bu tio n of tr ee s/ ha ;m os tly in co as ta la nd in la nd sa va nn ah . Sh ru b la nd an d th ic ke t A re as do m in at ed by de ns e sh ru bs an d th ic ke ts as so ci at ed w ith gr as sl an d. W at er bo dy R iv er s, po nd s, da m s, du go ut s, la ke s, an d la go on s – an yt hi ng w ith w at er su rf ac e. W et la nd M ar sh la nd s an d sw am ps . V eg et at io n is m os tly m an gr ov es an d se dg es . O pe n la nd L an d su rf ac es la ck in g an y fo rm of ve ge ta tio n co ve r, ei th er na tu ra lo r m an m ad e. B ui lt- up ar ea s Su rf ac es m od ifi ed by hu m an co ns tr uc tio n: e. g. ,c om m un iti es ,r oa ds ,a ir fie ld s, et c. Sa va nn ah w oo dl an d A re as in th e sa va nn ah re gi on w ith a hi gh de ns ity of tr ee s (> 15 0 tr ee s/ ha ). It al so in cl ud es pr ot ec te d ar ea s, su ch as M ol e N at io na lP ar k. T ab le 4. L an d us e ca te go ri es an d la nd co ve r di st ri bu tio ns in th e ec or eg io ns of G ha na . Su da n sa va nn ah G ui ne a sa va nn ah T ra ns iti on zo ne D ec id uo us fo re st R ai n fo re st W et ev er gr ee n C oa st al sa va nn ah 20 00 19 90 20 00 19 90 20 00 19 90 20 00 19 90 20 00 19 90 20 00 19 90 20 00 19 90 A gr ic ul tu ra l A gr ic ul tu ra l la nd 14 35 .7 8 24 11 .7 8 25 42 2. 26 21 24 6. 11 49 37 .1 0 89 26 .1 1 83 46 .2 7 68 73 .7 1 25 3. 28 32 9. 37 10 6. 90 18 0. 70 47 02 .5 1 32 79 .4 3 Fo re st ga rd en 29 5. 80 83 2. 65 32 21 1. 31 32 18 0. 90 14 02 8. 67 80 30 .1 8 29 44 5. 91 32 32 3. 12 65 04 .5 1 47 98 .3 1 37 39 .7 5 34 90 .9 8 34 40 .1 0 43 90 .2 7 Fo re st pl an ta tio n C lo se d fo re st 0. 00 0. 00 0. 00 0. 00 66 .4 2 66 .4 2 81 76 .4 5 81 76 .4 5 35 83 .9 2 35 83 .9 2 23 26 .8 6 23 26 .8 6 0. 00 0. 00 O pe n fo re st 0. 00 0. 00 0. 00 0. 00 14 99 .6 2 26 21 .5 5 79 61 .5 7 68 59 .0 9 13 59 .5 7 14 28 .8 5 71 7. 47 12 53 .0 7 7. 19 86 .2 2 Pl an ta tio n 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 22 75 .5 7 29 23 .2 3 48 1. 30 11 30 .3 8 35 4. 23 80 3. 08 9. 78 79 7. 27 W oo dl an d an d gr as s G ra ss la nd w ith tr ee s 77 0. 00 7. 27 50 65 .8 7 96 35 .6 6 19 61 .4 2 32 27 .6 7 31 96 .4 4 26 58 .9 2 0. 00 0. 00 0. 00 0. 00 21 42 .5 7 88 1. 45 Sa va nn ah w oo dl an d 32 .0 5 28 .2 5 20 46 .1 9 14 09 .1 3 56 1. 02 51 3. 21 20 6. 58 22 2. 31 0. 00 0. 00 0. 00 0. 00 7. 72 46 .5 4 Se ttl em en ta nd ba re la nd B ui lt – up ar ea s 33 .6 3 22 .5 9 12 84 .4 6 90 1. 99 26 8. 06 71 .1 0 29 90 .8 5 35 26 .6 1 20 .0 0 20 .5 4 11 0. 40 12 8. 90 28 05 .2 9 33 64 .0 7 O pe nl an d 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 12 6. 05 41 85 .4 8 0. 00 0. 00 0. 00 0. 00 52 4. 12 22 .4 6 W at er W at er bo dy 0. 99 0. 34 56 .4 6 38 .2 4 54 .6 2 37 .9 6 54 .8 8 37 .6 8 0. 70 0. 17 0. 33 0. 45 56 .3 2 3. 41 W et la nd 0. 00 0. 00 52 3. 14 40 8. 40 69 .3 0 12 8. 44 19 9. 28 94 .9 2 0. 00 0. 00 46 6. 53 48 .1 7 81 0. 16 12 9. 89 O th er Sh ru bl an d an d T hi ck et 0. 00 0. 00 0. 00 0. 00 12 64 7. 55 69 67 .7 2 27 58 0. 64 27 17 7. 61 0. 00 0. 00 0. 00 0. 00 88 04 .9 5 10 06 7. 24 456 Journal of Disaster ResearchVol.9 No.4, 2014 Land Use and Landscape Structural Changes in the Ecoregions of Ghana Fig. 2. Land cover distribution and changes from 1990 to 2000 in the Guinea savannah zone. nah woodlands in the Guinea savannah zone (Fig. 2) and with shrubland and thicket in the transition zone (Fig. 7). Though agricultural land area had a net decrease, the colossal increase in forest garden means that arable lands in all the ecoregions had a net increase. 3.1.2. Distribution of Forest and Plantation Land Cover The forest and plantation comprise open forest, closed forest, and plantation LCTs. Considering their composi- tion in all the ecoregions of Ghana, there was a net de- crease in closed forest, open forest and plantation. Open forest was not found in either the Sudan or Guinea savan- nah ecoregions. Deciduous forest ecoregions was domi- nated by open forest cover from 1990 to 2000; followed by the rainforest, wet evergreen, and transition zones. Interestingly, a portion of open forest was found in the coastal savannah ecoregion, though this was the least open forest area recorded. In all the ecoregions of Ghana where open forest was found, deforestation occurred in the open forest LCT with the highest open-forest loss taking place in the deciduous forest followed by the transition, wet ev- ergreen, and coastal savannah zones. The observed decrease in open forest was because it was replaced mainly by forest garden cover in the decid- uous forest (Fig. 3), rainforest (Fig. 4), and wet evergreen ecoregions (Fig. 5), and by agricultural land in the decid- uous forest region (Fig. 3). Plantation was largely not found in the Sudan savannah, Guinea savannah and transition zone ecoregions. In both study years, plantation LUT in rainforest was found to be larger than the evergreen forest and coastal savannah. In Ghana, about half of the forest plantation areas are found in the deciduous forest. Similar to open forest, there was a substantial decrease (deforestation) in plantation LCT in all the constituent ecoregions. However, the highest Fig. 3. Land cover distribution and changes in the deciduous forest zone for 1990 to 2000. plantation cover loss took place in the coastal savannah, followed by rainforest and wet evergreen. The deciduous forest had the least plantation cover. The decreased plantation cover was replaced by forest garden in the rainforest (Fig. 4) and by closed and open forest in the deciduous forest (Fig. 3). Closed forest was found to be missing in the entire three savannah ecoregions: namely, the Sudan, Guinea and coastal savannahs. Apart from these ecoregions in which closed forest was not found, in the remaining ecoregions, there was loss of closed forest LUT. Closed forest in both study periods was most dominant in the deciduous forest, followed by the rainforest. The lowest closed forest area in both study periods was observed in the transition zone. There was reduction (deforestation) in the closed forest in all the ecoregions except in the transition zone, where the closed forest area was maintained at 66.42 km2 (Table 4). All three forest types were most dominant in the decidu- ous forest followed by rainforest and wet evergreen. The decline in closed forest was replaced by agricul- tural land and forest garden in the deciduous forest ecore- gion (Fig. 3); open land and forest garden in the rainfor- est ecoregion (Fig. 4); and forest garden, open forest, and plantation in the wet evergreen ecoregion (Fig. 5). 3.1.3. Distribution of Settlement and Bare Land The settlement and bare land categories include built- up areas and open land. Built-up areas were found in all ecoregions. However, the true amount of built-up area is not accounted for in all the ecoregions. This is because of the 30 m image resolution used for the classification, which made it difficult to see most settlements under tree canopy, particularly in the rainforest where most of the settlements are buried under closed canopy cover. Urban sprawl occurred in the period studied and was highly as- sociated with decreases in deciduous forest and coastal Journal of Disaster ResearchVol.9 No.4, 2014 457 Antwi, E. K. et al. Fig. 4. Land cover distribution and changes in the rainforest zone of ghana from 1990 to 2000. Fig. 5. Land cover distribution and changes in the wet ever- green zone of ghana from 1990 to 2000. savannah. In these zones, urbanization rates are high, and they host the most densely populated towns, including the first and second largest cities of Ghana. The deciduous forest ecoregion and the coastal savannah had increased in built-up areas. The Guinea savannah ecoregion also experienced notable urban sprawl. The only LCT that increased in all the ecoregions during the study was the built-up. Even the built-up area that increased the least in any of the ecoregions amounted to more than twice the area occupied in 1990. Open land was found only in the deciduous and coastal savannah. The largest area (524.12 km2) occupied by open land was in the coastal savannah in 2000 followed by 22.46 km2 (Table 4) in 1990. In the coastal savannah, open land increased remarkably though the deciduous for- est experienced a reduction in open land area. The increase in built-up areas was taken from the dif- ferent LCTs across the nation. Figs. 6a and 6c show a notable increase in built-up areas in the coastal savannah where Accra, the capital city, is located. Fig. 6. The coastal savannah zone of Ghana for the year 1990 and 2000. (a) Grassland with trees increased within the coastal savannah zone of Ghana. (b) Forest garden in- creased within the coastal savannah zone. (c) Built-up and open areas increased for the coastal savannah zone. (d) Wet- lands and water bodies increased for the coastal savannah zone. 3.1.4. Distribution of Woodland and Grasses The woodland and grasses categories include grassland with trees and savannah woodland. Grassland with trees was not found in the rainforest and wet evergreen forest ecoregions. In both study periods, grassland with trees was highest in the Guinea savannah, particularly in 1990. Grassland with trees decreased in the Guinea savannah and transition zones, the highest decrease occurred in the Guinea savannah. On the other hand, the Sudan savan- nah, coastal savannah, and deciduous forest experienced an increase in grassland with trees. The highest increase occurred in the Sudan savannah. The grassland with trees, which had decreased by the end of the study in 2000, was replaced with savannah woodlands and agricultural land in the Guinea savannah (Fig. 2) and with agricultural land and forest garden in the transition zone (Fig. 7). There was no savannah woodland in the rainforest and wet evergreen. Similar to the grassland with trees, sa- vannah woodland was the most dominant in the Guinea savannah ecoregion in both periods studied; however, the year 2000 had the highest increase. The savannah wood- land cover decreased in all the component ecoregions ex- cept for Guinea savannah. The highest decrease occurred in the transition zone, with coastal savannah and transition zone undergoing similar reductions in area. The colossal increase in savannah woodland in the Guinea savannah ecoregion was large enough to cause an increase in the savannah woodland. The increase in savannah woodland was taken from grassland with trees and agricultural land in the Guinea savannah (Fig. 2) and from wetland in the Sudan savannah (Fig. 8). Shrubland and thicket was found only in the transition zone, deciduous forest, and coastal savannah. The highest increase in shrubland and thicket occurred in the transi- 458 Journal of Disaster ResearchVol.9 No.4, 2014 Land Use and Landscape Structural Changes in the Ecoregions of Ghana Fig. 7. Land cover changes and distributions in the transi- tion zone for 1990 to 2000. Fig. 8. Land cover distribution and changes in the sudan savannah region for 1990 to 2000. tion zone, followed by the deciduous forest. Shrubland and thicket was very dominant in the deciduous forest, particularly in 2000. 3.1.5. Distribution of Water Bodies and Wetlands The water bodies and wetlands categories have mainly areas occupied by water and wetlands. One of three LCTs found in all seven ecoregions is water bodies. Though slightly larger area was occupied by water bodies in the Guinea savannah, the distribution of water bodies in the transition zone, deciduous forest, and coastal savannah (in 2000 only) was very similar (Table 4). Though there was a decrease in water bodies in four of the ecoregions (wet evergreen, deciduous forest, transition zones, Guinea savannah), there was a general increase in area of water bodies. The increase in water bodies in the coastal savan- nah was the highest during the study, though a marginal increase was observed in the rainforest and Sudan savan- nah. Wetland was also found in all the ecoregions except in the Sudan savannah and rainforest ecoregions. The area occupied by wetlands was notably dominant in the 1990s in the Guinea savannah, coastal savannah, and transition zone. We should note that wetland is one of the three LCTs that decreased in all the ecoregions that have it. A large decrease occurred in the Guinea savannah and the transition zone. The observed increase in water bodies is found mostly in the coastal savannah ecoregion and along the Volta River (Fig. 6). The wetlands that decreased in the Su- dan savannah were often replaced by savannah woodland (Fig. 8). Though water bodies increased in the coastal sa- vannah. Figs. 6a and 6d show a notable increase in wet- land. 3.2. Landscape Structure Analysis of Different Ecoregions of Ghana The landscape metrics calculated revealed a more frag- mented landscape over the study years in all the ecore- gions of the country. The mean patch size (MPS), an in- dicator of the grain of the landscape [21], decreased by half within the 10 years of the study. MPS, which is one of the measurements of fragmentation, was generally re- duced consistently from 1990 to 2000 in all the ecore- gions. The highest MPS value, which was recorded in 1990 (66.9 km2), was nearly halved (39.6 km2) by 2000 (Table 5). The rainforest ecoregion recorded the most prominent reduction in MPS (46726.7 km2). On the other hand, the coastal savannah had the least reduction (8706.9 km2) in MPS. Another measure of fragmentation that also corrob- orates with the MPS is the NumP. However, in NumP, in- crements indicate a rise in fragmentation and vice versa. Similar trends of increased fragmentation were expected in this metric, and such was confirmed. For example, coastal savannah increased in NumP from 375 in 1990 to 532 in 2000 and rainforest from 107 to 248 (Table 5). Fragmentation in this context means the breaking up of habitat, ecosystems, or LCTs into smaller parcels [20]. Because larger patches tend to be more complex than smaller patches, fragmentation has the effect of determin- ing patch complexity independent of its size. Generally, all the patches that were calculated in the ecoregions for both years can be said to be complex in the sense that an MSI value of 1 indicates a square (in the case of grids) or a circle (in the case of polygons); any value lower or higher than 1 is an indication of simplicity or complexity, respectively [21]. The MSI estimated for ecoregions in both periods of study ranges from 1.6 in the year 2000 in the Sudan savannah to 2.5 in the year 1990 in the transition zones (Table 5). Thus, habitat types in the Sudan savannah in 2000 were the simplest, whereas those in the transition zone in 1990 were the most com- plex. Generally, MSI decreased in all ecoregions except for rainforest and wet evergreen. The highest decrease in MSI occurred in the transition zone, followed by the deciduous forest, with the rest having the same level of Journal of Disaster ResearchVol.9 No.4, 2014 459 Antwi, E. K. et al. T ab le 5. C om po si te m et ri cs sh ow in g la nd sc ap e st ru ct ur e ch ar ac te ri st ic s in th e ec or eg io ns of G ha na . L an ds ca pe m et ri cs C oa st al sa va nn ah D ec id uo us fo re st G ui ne a sa va nn ah R ai nf or es t Su da n sa va nn ah T ra ns iti on W et ev er gr ee n 19 90 20 00 19 90 20 00 19 90 20 00 19 90 20 00 19 90 20 00 19 90 20 00 19 90 20 00 SD I 1. 8 1. 7 1. 7 1. 8 1. 5 1. 4 1. 4 1. 3 1. 1 1. 2 1. 6 1. 5 1. 5 1. 3 SE I 0. 7 0. 7 0. 7 0. 7 0. 7 0. 7 0. 7 0. 6 0. 6 0. 7 0. 7 0. 6 0. 7 0. 6 M SI 2. 1 1. 8 2. 2 1. 7 2. 1 1. 8 1. 8 2. 1 1. 9 1. 6 2. 5 1. 8 2 2. 2 M PA R 35 .5 33 .9 12 45 .6 14 .1 33 .4 8. 7 53 .7 7. 4 49 .2 15 .3 24 .2 34 .5 18 .4 E D 3. 6 3. 6 2. 8 2. 6 2. 2 2. 6 2 2. 8 2. 6 2. 9 2. 6 2. 3 3. 1 3. 9 M PS 29 66 1. 8 20 95 4. 9 65 67 7. 8 35 36 3. 8 77 32 0. 2 48 47 8. 4 82 26 4. 6 35 53 7. 9 57 40 9. 1 17 33 5. 9 84 23 3 61 88 2. 7 40 50 1. 2 29 29 8. 3 N um P 37 5 53 2 11 46 21 28 15 35 24 48 10 7 24 8 50 16 5 37 7 51 3 20 6 28 5 T ab le 6. T ra ns iti on m at ri x of al ll an d co ve r ch an ge s in G ha na fr om 19 90 to 20 00 . A ct ua la re a is sh ow n in sq ua re ki lo m et er s an d pe rc en ta ge in re la tio n to th e to ta la re a of th e cl as s in 19 90 . L an d co ve r in 20 00 Landcoverin1990 A gr ic ul tu - ra ll an d Fo re st ga rd en Pl an ta tio n C lo se d fo re st O pe n fo re st G ra ss la nd w ith tr ee s Sh ru b la nd an d th ic ke t W at er bo dy W et la nd O pe n la nd B ui lt- up ar ea s Sa va nn ah w oo dl an d To ta l A gr ic ul tu ra l la nd 14 70 9. 21 (3 4% ) 15 06 0. 85 (3 4. 8% ) 46 6. 99 (1 .1 % ) 72 .6 1 (0 .2 % ) 66 .4 5 (0 .2 % ) 40 21 .6 6 (9 .3 % ) 48 4. 85 (1 .1 % ) 20 7. 29 (0 .5 % ) 44 .6 6 (0 .1 % ) 37 .7 4 (0 .1 % ) 56 3. 77 (1 .3 % ) 75 11 .1 3 (1 7. 4% ) 43 24 7. 21 (1 8. 1) Fo re st ga rd en 18 47 5. 57 (2 1. 5% ) 42 46 9. 28 (4 9. 4% ) 71 22 .5 2 (8 .3 % ) 81 7. 03 (0 .9 % ) 63 5. 02 (0 .7 % ) 46 63 .4 1 (5 .4 % ) 79 1. 7 (0 .9 % ) 18 5. 79 (0 .2 % ) 90 .9 6 (0 .1 % ) 42 .4 8 (0 .0 % ) 66 1. 9 (0 .8 % ) 10 09 0. 77 (1 1. 7% ) 86 04 6. 41 (3 6. 0) Pl an ta tio n 71 5. 34 (4 .5 % ) 87 90 .6 4 (5 5. 1% ) 54 61 .3 3 (3 4. 2% ) 38 0. 96 (2 .4 % ) 28 9. 55 (1 .8 % ) 16 0. 23 (1 .0 % ) 1. 9 0. 0% ) 51 .9 9 (0 .3 % ) 24 .9 5 (0 .2 % ) 17 .5 5 (0 .1 % ) 55 .7 (0 .3 % ) 7. 2 (0 .0 % ) 15 95 7. 34 (6 .7 ) C lo se d fo re st 30 .0 9 (0 .2 % ) 96 9. 45 (7 .9 % ) 39 1. 47 (3 .2 % ) 95 28 .2 6 (7 7. 8% ) 12 35 .4 8 (1 0. 1% ) 14 .2 2 (0 .1 % ) 9. 82 (0 .1 % ) 8. 1 (0 .1 % ) 3. 19 (0 .0 % ) 2. 47 (0 .0 % ) 56 .2 5 (0 .5 % ) 12 24 8. 79 (5 .1 ) O pe n fo re st 44 6. 04 (7 .9 % ) 25 89 .0 9 (4 5. 8% ) 59 8. 15 (1 0. 6% ) 68 0. 89 (1 2% ) 57 2. 8 (1 0. 1% ) 49 .7 9 (0 .9 % ) 29 3. 38 (5 .2 % ) 14 .1 6 (0 .3 % ) 8. 87 (0 .2 % ) 0. 76 (0 .0 % ) 7. 79 (0 .1 % ) 39 2. 25 (6 .9 % ) 56 53 .9 6 (2 .4 ) G ra ss la nd w ith tr ee s 31 22 .6 (1 9. 0% ) 46 87 .2 (2 8. 6% ) 3. 13 (0 .0 % ) 29 .9 3 (0 .2 % ) 33 .6 3 (0 .2 % ) 13 71 .4 9 (8 .4 % ) 25 0. 32 (1 .5 % ) 74 .9 7 (0 .5 % ) 52 .4 2 (0 .3 % ) 0 (0 .0 % ) 75 .1 7 (0 .5 % ) 67 10 .1 1 (4 0. 9% ) 16 41 0. 97 (6 .9 ) Sh ru b la nd an d th ic ke t 38 2. 9 (1 7. 3% ) 42 5. 88 (1 9. 2% ) 18 .6 3 (0 .8 % ) 14 4. 07 (6 .5 % ) 33 0. 97 (1 4. 9% ) 83 5. 25 (3 7. 6% ) 4. 55 (0 .2 % ) 0. 74 (0 .0 % ) 21 .1 5 (1 .0 % ) 55 .2 9 (2 .5 % ) 22 19 .4 3 (0 .9 ) W at er bo dy 37 3. 96 (4 .7 % ) 44 6. 25 (5 .6 % ) 4. 16 (0 .1 % ) 0. 67 (0 .0 % ) 0. 2 (0 .0 % ) 21 6. 29 (2 .7 % ) 3. 98 (0 .0 % ) 66 60 .7 5 (8 2. 9% ) 67 .6 9 (0 .8 % ) 66 .4 8 (0 .8 % ) 16 .5 (0 .2 % ) 17 8. 84 (2 .2 % ) 80 35 .7 9 (3 .4 ) W et la nd 74 0. 3 (1 7. 6% ) 13 83 .7 8 (3 2. 9% ) 44 .3 (1 .1 % ) 6. 12 (0 .1 % ) 6. 63 (0 .2 % ) 36 9. 98 (8 .8 % ) 41 .9 3 (1 .0 % ) 14 1. 77 (3 .4 % ) 28 2. 97 (6 .7 % ) 8. 05 (0 .2 % ) 15 .0 1 (0 .4 % ) 11 67 .0 8 (2 7. 7% ) 42 07 .9 3 (1 .8 ) O pe n la nd 1. 94 (1 .6 % ) 42 (3 5. 5% ) 27 .2 3 (2 3% ) 5. 98 (5 .1 % ) 0. 4 (0 .3 % ) 0. 02 (0 .0 % ) 13 .0 8 (1 1. 1% ) 4. 69 (4 .0 % ) 22 .9 3 (1 9. 4% ) 11 8. 26 (0 .0 ) B ui lt- up ar ea s 72 .2 1 (8 .9 % ) 60 .3 3 (7 .4 % ) 31 .2 7 (3 .9 % ) 0. 9 (0 .1 % ) 3. 05 (0 .4 % ) 10 .6 8 (1 .3 % ) 12 .7 4 (1 .6 % ) 6. 09 (0 .8 % ) 7. 2 (0 .9 % ) 33 .1 2 (4 .1 % ) 56 4. 93 (6 9. 8% ) 7. 31 (0 .9 % ) 80 9. 82 (0 .3 ) Sa va nn ah w oo dl an d 61 33 .9 4 (1 3. 9% ) 12 74 1. 33 (2 8. 8% ) 3. 1 (0 .0 % ) 3. 37 (0 .0 % ) 13 4. 01 (0 .3 % ) 19 27 .5 8 (4 .4 % ) 12 7. 66 (0 .3 % ) 15 6. 81 (0 .4 % ) 69 .6 9 (0 .2 % ) 1. 86 (0 .0 % ) 79 .3 3 (0 .2 % ) 22 83 3. 81 (5 1. 6% ) 44 21 2. 48 (1 8. 5) To ta l 45 20 4. 1 (1 8. 9) 89 66 6. 1 (3 7. 5) 14 15 3. 65 (5 .9 ) 11 54 5. 4 (4 .8 ) 31 20 .8 9 (1 .3 ) 13 13 6. 3 (5 .5 ) 28 53 .5 3 (1 .2 ) 75 12 .6 7 (3 .1 ) 65 0. 17 (0 .3 ) 22 4. 31 (0 .1 ) 20 68 .4 1 (0 .9 ) 49 03 2. 97 (2 0. 5) 23 91 68 .4 (1 00 ) 460 Journal of Disaster ResearchVol.9 No.4, 2014 Land Use and Landscape Structural Changes in the Ecoregions of Ghana decrease in habitat shape complexity. Two ecoregions in- creased in complexity – rainforest and wet evergreen – which had MSI values of up to 2.1 and 2.2 by the year 2000 respectively (Table 5). With the exception of wet evergreen and rainforest, habitat complexity was higher in ecoregions that had larger mean patch sizes. Another important feature for any landscape is its ex- tent of heterogeneity, which can be quantified by the Shannon Diversity Index (SDI) and the Shannon Even- ness Index (SEI). The value of SDI and SEI is 0 when there is only one patch in a landscape, which is an indi- cation of evenness in the case of SDI and of dominance in the case of SEI. The coastal savannah and the decid- uous forest ecoregion of the country showed the highest values of SDI, with values between 1.7 and 1.8 (Table 5) between 1990 and 2000. All other ecoregions exhibited a milder form of diversity, with SDI values ranging from 1.6 in the transition ecoregion for the year 1990 to 1.1 for Sudan savannah for the same year (Table 5). SEI val- ues, which also complement the SDI values, remained at 0.7 for coastal savannah, deciduous forest, and Guinea sa- vannah in both 1990 and 2000 (Table 5). Table 6 shows transition in all LCCs across Ghana from 1990 to 2000. 4. Discussion 4.1. Land Use Distribution and Land Cover Changes in the Ecoregions of Ghana The observed LC transformations in the form of defor- estation, urban sprawl, agricultural expansion, and water resources losses in the ecoregions of Ghana have impor- tant implications for sustainable land use management. Specifically, reductions in forest (closed and open), and savannah woodland (closed and open) with accompany- ing increments in settlement/built-up areas (throughout the ecoregions) and grassland/unharnessed farmlands in the Sudan savannah ecoregion have been observed else- where [13]. 4.1.1. Land Cover Changes and Deforestation in the Ecoregions of Ghana Plantation, closed forest and open forest decreased con- siderably in the evergreen, deciduous forest and rainfor- est ecological zones. Though most of the forest area was maintained, substantial portions of the closed and open forest have been converted to non-forest land (Figs. 3- 5) such as built-ups, farmlands, and open fields at the fringes [22]. Particularly, farming at the fringes of for- est while increasing poor people’s access to natural re- sources, has generally been observed as a common prac- tice that influences LUC in most forest ecoregions in de- veloping countries [22-25]. Ghana’s 1948 Forest Policy designated all areas outside permanent forests as areas available for maximum utiliza- tion or authorized conversion to other LCTs [26]. This policy led to the expansion of farms, especially the cocoa- annuals-forest mosaic, plantations and forest gardens in the forest ecoregions. The implementation of the Struc- tural Adjustment Program from 1983 to the early 1990’s which encouraged the expansion of the timber industry and contributed to further loss of forest cover [27, 28], see Figs. 3-5. However, the promulgation of the 1994 Ghana Forestry and Wildlife Policy restricted LU conversions to land outside protected areas [29]. This limited subsequent land conversions outside the protected reserves in the rain- forest and wet evergreen forest. Currently, restriction on the conversion of forest in protected areas has been rein- forced by the amended forest and wildlife policy [28]. 4.1.2. Land Cover Changes and Urban Sprawl in the Ecoregions of Ghana Urban sprawl is a multidimensional concept that often involves outward spreading of cities and their suburbs. This is often attributed to natural increase and rural-urban migration [30]. The ecological significance of LC change in urban ar- eas cannot be examined separately from forest land en- vironments because the two are directly and indirectly related with synergistic effects on the environment. LU type in an area is determined by the natural and socio- economic factors as well as the temporal and spatial use which people assign to the land [31]. The observed LCCs between the study periods highlight the importance of ur- ban sprawl and associated population growth with regard to urban and forest lands. Urban sprawl is an issue of con- cern in many developing nations; it often has a damaging effect on natural resources and infrastructure [32]. The LCC changes between 1990 and 2000 in all the ecore- gions show expansion in the built-up areas. This was of- ten associated with the conversion of forest into arable, built-up land or direct conversion to open land. According to [22], vegetation cover has been decreasing in Ghana, giving way to other LUs, such as open and built-up lands, for the past three decades. Ghana is becoming more ur- banized at a fast rate; about one-third of Ghanaians are now living in towns and cities [33]. The three most pop- ulous cities, Accra, Kumasi and Takoradi, located in the in the coastal savannah zone, moist deciduous forest and in the wet evergreen zone respectively, are the biggest culprits in turning vegetated areas into built-up and open areas. Though significant, actual observed increases in built-up areas could not be accounted for in our study due to the resolution of the images used (30 m) which could not account for settlements covered by tree canopy. For instance, the classification output could not show built- up areas in the rainforest zone because of the tree canopy cover. 4.1.3. Land Cover Changes and Agricultural Expan- sion in the Ecoregions of Ghana While there was a net decrease in agricultural land in most of the ecoregions, the colossal expansion of agricul- tural land particularly forest gardens in the Guinea savan- nah and deciduous forest zone, both of which account for over 60% of the LCTs in Ghana, led to an expansion in Journal of Disaster ResearchVol.9 No.4, 2014 461 Antwi, E. K. et al. agricultural land. Agriculture is arguably the most domi- nant LU in Ghana, given that about 41.2% of the econom- ically active population of the nation is engaged in it [34]. In most ecoregions of Ghana, increases in yield are at- tributed to agricultural land use expansion. For instance, land accounts for 65% of the total agriculture value added in Northern Ghana [35] (located within the Guinea Savan- nah ecoregion), which indicates a higher land-to-labor ra- tio than the rest of the country. This continuing process of swift anthropogenic LUC could affect numerous wildlife species, especially species that require large areas for sur- vival. Such open lands as roads, gaps, and ploughed lands act as barriers to movement for many animal species [36- 38]. 4.1.4. Impacts of Land Use Change on Water Re- sources in the Ecoregions of Ghana Generally, the monitored landscape transformation recorded a decrease in water bodies in the wet evergreen, deciduous forest, transition zones and Guinea savannah. Also, all the ecoregions that had wetlands particularly in the guinea savannah zone decreased. Decreases or losses in wetlands and water bodies could have serious impli- cations for well-being. This is because water bodies and wetlands play very important role in landscape function- ing particularly for humans and other life forms. For instance, wetlands are critical for controlling floods, re- moval of pollutants from water, groundwater recharge, as well as providing habitat for wildlife, and serve im- portant recreational and cultural functions. The growth of cities particularly in new developing nations has of- ten led to encroachment on watercourses as new build- ings are constructed, either legally or illegally. The en- croachment of developments on wetlands and waterways has been a major challenge for most of Ghana’s major ur- ban settlements. This is often associated with increased impervious surfaces within urban catchments, changing the hydrology and geomorphology of streams [39]. The yearly occurrence of floods in Accra, Ghana’s capital city can partly be blamed on the conversion and destruction of wetlands and poor drainage management. Aside from reduction in wetlands and water bodies, wa- ter quality is most vulnerable in the urban and industrial- ized areas mostly in the southern part of Ghana [40]. Log- ging (lumbering) and mining are the principal causes of the changing landscape and water resources in the south- ern part of the country [41]. Particularly, the widespread activities of small scale illegal miners who employ meth- ods that result in vegetation removal, erosion, and siltation and sedimentation of river bodies [42] have been a ma- jor culprit. In addition, both large- and small-scale min- ers, as well as illegal chain-saw operators, are threaten- ing several forest reserves with mineral resources under- neath [41]. The role of mining in the land use changes is particularly damaging in nature and regardless of the method used and resources available, it affects land cover at the site of extraction, leaving behind multiple damages that stretch over a wide range of land. Higher amounts of mercury, arsenic, and other poisonous substances than are acceptable byWHO standards are found in most water bodies around the mining areas in Ghana [43, 44]. 4.2. Landscape Configuration and Composition in the Ecoregions of Ghana Demand for land and land-based natural resources to support competing livelihoods and developmental activi- ties in the different ecoregions have changed the structure of the Ghanaian landscape [45]. These changes occur on different scales and include modifications in size, shape, and composition and spatial configurations of landscape features which affect habitat fragmentation, habitat rich- ness, habitat complexity and diversity in the ecoregions of Ghana. Habitat fragmentations are often accompanied by habi- tat loss, resulting in a major impact on the regional survival of plant species. With habitat sizes becoming smaller across the ecoregions, plant species that prefer in- terior habitat conditions could become more susceptible to extinction though some level of fragmentation is actu- ally preferred by more generalist species. If the area of individual habitats is reduced, such species become vul- nerable to external influences, hence affecting the survival potential of populations in these patches [46]. The decreasedMPS, coupled with an increased number of patches, could sometimes lead to possible decreases in population sizes and reduction in habitat diversity [47]. This is particularly evident in the decrease in diversity (according to the SDI) in all ecoregions except decidu- ous forest and Sudan savannah. On the other hand, MPSs generally reveal that habitat sizes in the closed forest and open forest have significantly decreased, indicating the fragmentation of the forest LC. Smaller fragments formed may not contain interior habitat, but they support smaller species’ populations, which tend to be vulnerable to ex- tinction. Observed habitat shape in the Sudan savan- nah in 2000 became the simplest in all the periods and ecoregions studied. The formation of such simple patches could create variability in habitat opportunity since small patches or simple patches often do not provide the same habitat opportunity as larger patches, particularly for or- ganisms that prefer interior habitat conditions. In all ecoregions, except wet evergreen and rainforest, patches in landscape that have larger mean patch sizes also have high shape complexity. This change in habitat irregularity stems from the fact that large habitats some- times show irregular shapes [48] and confirms the ob- servation that when patches join to become larger, they do not necessarily become simple [49]. The creation of straight patch edges particularly at the fringes of the for- est could be attributed to the restriction in forest depletion to outside protected areas and sacred community groves as required by the 1994 Ghana Forestry and Wildlife Pol- icy [29]. The SDI and SEI are measures of diversity in a commu- nity. The reductions in diversity for natural landscapes, such as moist evergreen forest, wet evergreen forest, tran- 462 Journal of Disaster ResearchVol.9 No.4, 2014 Land Use and Landscape Structural Changes in the Ecoregions of Ghana sition zones, and coastal and Guinea savannah zones, sug- gest a less varied landscape. The deciduous forest and Sudan savannah recorded high diversity and could suggest more diversity. It is not surprising that the moist evergreen forest, wet evergreen forest, transition zone, and coastal and Guinea savannah zones were less diverse because these are highly fragmented regions in Ghana as they host high number of socio-economic developments involving urban development and mining activities. Increased frag- mentation in the ecoregions of Ghana was associated with loss of habitat diversity and population [47, 50, 51]. The general increase in built-up areas and agricultural land use expansion in almost all the ecoregions between 1990 and 2000 could be one of the main causes of fragmen- tation. Growing urban areas and intensive agricultural LU may narrow and separate the remaining wildlife habi- tats [20, 38]. Long absence of disturbance or severe disturbance has a depressing effect on diversity, though an intermediate level of disturbance in a landscape has been reported to enhance diversity [52]. In the Ghanaian landscape, distur- bance from rigorous deforestation, the turning of vegeta- tion into open lands, and increases in built-up areas over the 10 years of the study have had a depressing effect on landscape diversity in all ecoregions except for the decid- uous forest and Sudan savannah. Disturbances in the de- ciduous forest and Sudan savannah are indicated at an in- termediate level. The intermediate disturbance hypothesis noted above predicts that such intermediate disturbances will result in increased diversity [53]. 5. Conclusions LU type in any area is determined by the natural and socio-economic factors as well as the temporal and spa- tial uses assigned to the land [31]. In the ecoregions of Ghana, the LCCs reflect deep structural issues, involving LU policies that exact more resources than normal from natural landscapes. This LCCs assessment indicate that changes in LU and landscape structure are in the form of forest degradation, urban sprawl, agricultural expansion, and destruction of water resources in the ecoregions of Ghana. Although other similar studies have been carried out, this research introduces the first attempt to make use of tools available in GIS and remote sensing as well as landscape ecology to expand the pattern-process relation- ship occurring in Ghana to a country scale. Substantial areas of closed and open forest have been converted to non-forest land (semi-natural areas) on the fringes. An increase in built-up areas, farmlands, open fields, and grassland and an accompanying loss of forests was observed across the ecoregions of Ghana [54, 55, 13]. Instances of afforestation were very rare, occurring only in pockets in the rainforest. Demand for land and land- based resources to support competing livelihoods and de- velopmental activities in the different ecoregions owing to rising urban population have brought severe losses to for- est and agricultural lands, shrubs, barren land, and water bodies. In the southern and western regions of Ghana, large and small-scale mining activities and logging (lumbering) have been the principal causes of the changing landscape and water resources. Though logging remains one of the leading causes of forest degradation [56], forest loss in Ghana varies across ecoregions. Forest in Northern Ghana (Guinea and Sudan savannah) is lost to wild fires, charcoal production, wood fuel harvesting, and farming. In all ecoregions except wet evergreen and rainforest, patches in landscape that have larger mean patch sizes also have high shape complexity. The reduced habitat shape complexity in most ecoregions clearly indicates in- creased anthropogenic influences on the landscape since human-made edges tend to be more regular in shape. In most ecoregions except for wet evergreen and rainforest, the availability of unaffected core patch areas may how- ever not been reduced. Acknowledgements This research is carried out with the support of the CECAR Africa project funded by JSTS/JICA. References: [1] Food and Agriculture Organization (FAO), “Global forest resources assessment 2010,” Food and Agriculture Organization of the United Nations, Rome, 2010. [2] J. 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Ghazoul, “Implementing landscape fragmenta- tion as an indicator in the Swiss Monitoring System of Sustainable Development (MONET),” Journal of Environmental Management, Vol.88, No.4, pp. 737-751, 2008. 464 Journal of Disaster ResearchVol.9 No.4, 2014 Land Use and Landscape Structural Changes in the Ecoregions of Ghana Name: Effah Kwabena Antwi Affiliation: Assistant Professor, Integrated Research System for Sustainability Science (IR3S), The Univer- sity of Tokyo Visiting Fellow, United Nations University – In- stitute for Advanced Study of Sustainability Address: 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-8654, Japan Brief Career: 2001-2005 Researcher and MSc. Student, Department of General Ecology, Brandenburg University of Technology, Germany 2006-2009 Doctoral Candidate and Assistant Lecturer, Department of General Ecology, Brandenburg University of Technology, Germany 2008-2011 Teaching Assistant, Department of Geography, University of Ghana, Legon 2009-2011 JSPS Postdoctoral Fellow United Nations University-Institute for Advanced Study of Sustainability, Integrated Research System for Sustainability Science (IR3S), The University of Tokyo 2011 MSc. Degree, Institute of Sustainability and Peace, United Nations University 2011-2014 Project Assistant Professor, IR3S, University of Tokyo Visiting Fellow, United Nations University-Institute for Advanced Study of Sustainability Selected Publications: • “Assessing Landcover Changes from Coastal Tourism Development in Ghana: Evidence from the Kokrobite – Bortianor Coastline, Accra,” Civil and Environmental Research, Vol.6, No.6, 2014 (co-authored). • E. K. Antwi, R. Krawczynski, and G. Wiegleb, “Detecting the Effect of Disturbance on Habitat Diversity and Land Cover Change in a Post-Mining Area Using GIS,” Landscape and Urban Planning, Vol.87, pp. 22-32, 2008. •R. Krawczynski and E. K. Antwi, “Reversed Succession in Post-Mining Landscape and Possible Causes,” Biological Studies, Vol.39, pp. 48-58, Luckau, Germany, 2010. Academic Societies & Scientific Organizations: • Japan Society for the Promotion of Science (JSPS) • Member, Evaluation and Selection Committee, the MIDORI Prize for Biodiversity Name: John Boakye-Danquah Affiliation: Teaching Assistant, Department of Geography and Resource Development, University of Ghana Address: P.O.Box LG 59, Legon-Accra, Ghana Brief Career: 2007- B.A. Degree (Hons), University of Ghana 2008-2011 Teaching Assistant, Department of Geography, University of Ghana, Legon 2011- M.Sc. Degree, Institute of Sustainability and Peace, United Nations University Selected Publications: • J. Boakye-Danquah, “The Impact of Agriculture Land Use Change and Farm Management Practices on Soil Organic Carbon Sequestration Potential: The Case of Savannah Regions of Northern Ghana,” M.Sc. Dissertation, The United Nations University – Institute of Sustainability and Peace (UNU-ISP), July, 2013. Academic Societies & Scientific Organizations: • African Studies Association of Australasia and the Pacific (AFSAAP) Name: Stephen Boahen Asabere Affiliation: Ph.D. Student, Department of Geosceinces, Technische Universitaet Dresden Address: 01062 Dresden, Germany Brief Career: 2007-2008 Intern, Resource Management Support Center 2008-2009 Forestry Commission Ghana Academic Societies & Scientific Organizations: • International Association of Landscape Ecology, Germany (IALE) • African Association of Remote Sensing of the Environment (AARSE) • International Association of Bamboo and Rattan (IABR) Name: Gerald A. B. Yiran Affiliation: Assitant Lecturer, Department of Geography and Resource Development, University of Ghana Address: P.O.BOX LG 59, Legon-Accra, Ghana Brief Career: 2001-2009 Senior Cartographer, Department of Geography and Resource Development, University of Ghana 2009-present Assistant Lecturer, Department of Geography and Resource Development, University of Ghana 2011-present Ph.D. Student, Department of Geography and Resource Development, University of Ghana Selected Publications: • E. A. Gyasi, G. Kranjac-Berisavljevic, M. Fosu, A. M. Mensah, G. Yiran, and I. Fuseini, “Managing Threats and Opportunities of Urbanisation for Urban and Peri-urban Agriculture in Tamale, Ghana,” In: “The Security of Water, Food, Energy and Liveability of Cities: Challenges and Opportunities for Peri-Urban Futures,” B. Maheshwari, R. Purohit, H. Malano, V. P. Singh, and P. Amerasinghe (Eds.), Springer, Vol.71, pp. 87-97, 2014. • G. A. B. Yiran, J. M. Kusimi, and S. K. Kufogbe, “A synthesis of remote sensing and local knowledge approaches in land degradation assessment in the Bawku East District, Ghana,” International Journal of Applied Earth Observation and Geoinformation, Vol.14, pp. 204-213, 2012. • S. M. Yidana, G. A. B. Yiran, P. A. Sakyi, P. M. Nude, and B. Banoeng-Yakubo, “Groundwater Evolution in the Voltaian Basin, Ghana – An application of multivariate statistical analyses to hydrochemical data,” Natural Science Journal, Vol.3, No.10, pp. 837-854 , 2011. Academic Societies & Scientific Organizations: • Next Generation of Researchers, Africa (NGR) • Centre for Climate Change Economics and Policy (CCCEP), University of Leeds, UK Journal of Disaster ResearchVol.9 No.4, 2014 465 Antwi, E. K. et al. Name: Seyram Kofi Loh Affiliation: University of Ghana Office, Climate and Ecosys- tem Change Adaptation and Resilience Research (CECAR-AFRICA) Address: P.O.Box LG 59, Legon-Accra, Ghana Brief Career: 2009- Teaching/Research Assistant, College of Agriculture and Natural Resource Management, Kwame Nkrumah University of Science and Technology (KNUST) 2012- Remote Sensing Data Analyst, Centre for Remote Sensing and Geographic Information Services (CERSGIS), University of Ghana-Legon (www.cersgis.org) 2013- GIS Research Assistant, Climate and Ecosystem Change Adaptation and Resilience Research (CECAR-AFRICA), University of Ghana, Legon, Accra Academic Societies & Scientific Organizations: • Centre for Geospatial Analysis and Mapping (CEGAM) Name: Kwabena Gyekye Awere Affiliation: Lecturer, University of Ghana Address: P.O.BOX LG 57 Legon, Ghana Brief Career: Worked at EPA Ghana for six years as Programme Officer Completed Ph.D. in 2005 from St. Petersburg State University Joined University of Ghana in 2006 as Lecturer Selected Publications: • A. K. Gyekye, “An Assessment of Toxic in Urban Soils Using Garden Cress, (Lepidium sativum) in Vasileostrovsky Ostrov and Elagin Ostrov, Saint Petersburg, Russia,” Journal of Geography and Geology, Vol.5, No.4, Canadian Center of Science and Education, pp. 75-100, 2013. • A. K. Gyekye, “Environmental Change and Flooding in Accra, Ghana. Sacha Journal of Environmental Studies,” Vol.3, No.1, London, United Kingdom, pp. 65-80, 2013. • A. K. Gyekye, “Chemical characteristics of urban soils of Vasileostrovsky Ostrov and Elagin Ostrov, St Petersburg, Russia,” West Africa Applied Ecology, Vol.21, No.2, pp. 121-133, 2013. • A. K. Gyekye, J. M. Kusimi, and A. B. Yiran Gerald, “Geomorphological Processes and Landforms of the Coastal Environment,” In G. Owusu, S. Agyei-Mensah, P. W. K. Yankson, and E. M. Attua (Eds.), Selected Readings in Geography Reader, Accra Woeli Publishing Services, pp. 248-264, 2013. Academic Societies & Scientific Organizations: • Geographical Society of Ghana (GSG) Name: Felix K. Abagale Affiliation: Senior Lecturer, Faculty of Agriculture, Univer- sity for Development Studies Address: P.O.Box 1350, Tamale, Ghana Brief Career: Lecturing in the Faculty of Agriculture, with special interest in Agrometeorology and Environmental Management, with about 7 years of experience. He has several scientific reseacrh articles in the area of environment, waste, soil and water to his credit. Selected Publications: • F. K. Abagale, N. Kyei-Baffour, and E. Ofori, “Degradation of the Nasia River Basin in Northern Ghana,” Ghana Journal of Development Studies (GJDS), Vol.6, No.1, 2009. • K. Unami, T. Kawachi, G. Kranjac-Berisavljevic, F. K. Abagale, S. Maeda, and J. Takeuchi, “Case study: Hydraulic Modeling Of Runoff Processes In Ghanaian Inland Valleys,” Journal of Hydraulic Engineering, America Society of Civil Engineers (ASCE), Vol.135, No.7, pp. 539-553, 2009. Academic Societies & Scientific Organizations: • Ghana Science Association (GSA) Name: Kwabena Owusu Asubonteng Affiliation: Institute for Natural Resources in Africa (UNU- INRA), United Nations University Address: Annie Jiagge Road, University of Ghana Campus, Legon-Accra, Ghana Brief Career: 2007- Sustainable Land Management Project, Ghana 2009- APERL GIS Training and Research Centre, KNUST Sunyani Campus, Ghana 2011- Geo-Information Analyst, UNU-INRA, Ghana Selected Publications: • A. T. Koomson and K. O. Asubonteng, “Collaborative governance in extractive industries in Africa,” United Nations University Institute for Natural Resources in Africa, Accra, 2013. • D. Tutu-Benefoh, S. Oppong, K. O. Asubonteng, L. Addae-Wireko, and E. Acheampong, “Rapid assessment of non-market values of carbon sequestration services from rural community-actors in Ghana,” American Journal of Scientific and Industrial Research, 2010. Academic Societies & Scientific Organizations: • Ghana Institute of Professional Foresters (GIPF) 466 Journal of Disaster ResearchVol.9 No.4, 2014 Land Use and Landscape Structural Changes in the Ecoregions of Ghana Name: Emmanuel Morgan Attua Affiliation: Senior Lecturer, Department of Geography and Resource Development, University of Ghana Address: P. O. Box LG 59, Legon, Accra, Ghana Brief Career: 1996-2000 Research Assistant/Teacher Demonstrator, Department of Botany, University of Ghana, Legon 2000-2006 Lecturer, Department of Geography and Resource Development, University of Ghana, Legon 2007-present Senior Lecturer, Department of Geography and Resource Development, University of Ghana, Legon 2008 Commonwealth Visiting Scholar, University of Oxford, United Kingdom Selected Publications: • “Relating Land Use and Land Cover to Surface Water Quality in the Densu River basin, Ghana,” International Journal of River basin Management, Vol.12, No.1, pp. 57-68, March, 2014. • “Historical and future land cover change in a municipality of Ghana,” Earth Interactions, Vol.15, No.9, pp. 1-26, 2011. • “Rehabilitation of forest-savannas in Ghana: The impacts of land use, shade, and invasive species on tree recruitment,” Applied Geography, Vol.31, No.1, pp. 181-190, 2011. • “Sustainable land-use evaluation on steep landscapes and flood plains in the New Juaben district of Ghana: A GIS Approach,” Ghana Journal of Geography, Vol.1, pp. 115-134, 2009. Academic Societies & Scientific Organizations: • Ghana Geographical Association (GGA) • Ghana Science Association (GSA) • University Teachers Association, Ghana (UTAG) Name: Alex Barimah Owusu Affiliation: Lecturer, Department of Geography and Re- source Development, University of Ghana Address: LG 59, Legon, Ghana Brief Career: 2006-2009 Research Assistant, Department of Geography and Geoinformation Science; George Mason University, Fairfax VA, USA 2007-2008 Adjunct Professor, Department of Humanities and Social Sciences; Northern Virginia Community College, Alexandria VA, USA 2010-present Lecturer, Department of Geography and Resource Development; University of Ghana, Legon Selected Publications: • A. B. Owusu, “Measuring Desertification in continuum: Normalized Difference Vegetation Index-based Study in the Upper East Region, Ghana,” Journal of Civil and Environmental Research, Vol.3, No.12, pp. 157-170, 2013. • A. B. Owusu, S. Frimpong, and S. Abrokwah, “Analysis of the Spatial and Temporal Dynamics of Street Hawking: A Case Study of the Accra Metropolitan Area,” Journal of Geography and Geology, Vol.4, No.4, pp. 1-12, 2013. • A. B. Owusu, “Detecting and Quantifying Desertification in the Upper East Region, Ghana using Multi spatial and multi temporal Normalized Difference Vegetation Index,” Journal of Environment and Earth Science, Vol.3, No.10, pp. 62-78, 2013. • A. B. Owusu, C. Guido, and S. L. Beach, “Analysis of Desertification in the Upper East Region (UER), Ghana using remote sensing, field study and local knowledge,” Cartographica, pp. 1-24, 2013. Academic Societies & Scientific Organizations: • Association of American Geographers (AAG) • Ghana Geographical Association (GGA) • Borlaug Norman LEAP Fellowship Journal of Disaster ResearchVol.9 No.4, 2014 467